Wiens, J.A. 2002. Riverine landscapes: taking landscape into the water. Freshwater Biology 47:501516

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Wiens, J.A. 2002. Riverine landscapes taking
landscape into the water. Freshwater Biology
47501-516.
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River as element of landscape mosaic
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River linked with surroundings by boundary
dynamics
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River as internally heterogeneous landscapes
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Central Themes Relevant to Rivers

• Patches differ in quality
• Patch boundaries affect flows

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Central Themes Relevant to Rivers

• Patches differ in quality
• Patch boundaries affect flows
• Patch context matters
• Connectivity is critical
• Organisms are important
• Importance of scale
• Spatial and temporal

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Low-Relief Grasslands
(A)
mobile FPZ
constrained lowland FPZ
unconstrained headwater FPZ
mobile FPZ
armoured FPZ
mobile FPZ
unconstrained lowland FPZ
meandering FPZ
anabranch FPZ
Upland Forests
floodplain FPZ
constrained headwater FPZ
delta FPZ
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Low-Relief Grasslands
(A)
mobile FPZ
constrained lowland FPZ
unconstrained headwater FPZ
mobile FPZ
armoured FPZ
mobile FPZ
unconstrained lowland FPZ
meandering FPZ
anabranch FPZ
Upland Forests
floodplain FPZ
constrained headwater FPZ
delta FPZ
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• flow regime, flow history, flood pulse (long to
  short time scales)
• hydraulics
• local geology
• topography and slope
• substrate size/composition
• radiation (PAR, UVR)
• temperature
• oxygen
• inorganic nutrients
• other water chemistry

River Network

• Habitat Forming
• dams (e.g., beavers)
• retreats (e.g., caddisfly cases)
• species clumps (e.g., zebra mussels)
• biofilm
• bioturbation
• Density Patches
• reproduction/production
• dispersal/migration
• food/habitat aggregation
• Negative Interactions
• competition
• predation herbivory
• parasitism, disease

• Types
• terrestrial vegetation
• benthic and pelagic, micro-
• and filamentous algae
• vascular macrophytes
• detrital (leaves, woody debris)
• Mechanisms
• terrestrial abiotic disturbances
• aquatic abiotic disturbances
• radiation, nutrients, etc.
• biotic disturbance
• autotroph-autotroph interactions
• individual species requirements

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Orinoco River - Venezuela
Varzea