E

1
1
Phe
fMet
E
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
5?
3?
P
A
2
Peptide bond
1
2
fMet
Phe
Phe
fMet
E
E
U
A
C
A
A
A
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
5?
3?
5?
3?
P
A
P
A
3
Peptide bond
3
1
2
fMet
Phe
fMet
Phe
Phe
fMet
Movement of ribosome
E
E
U
A
C
A
A
A
U
A
C
A
A
A
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
5?
3?
5?
3?
5?
3?
P
A
P
A
P
A
E
4
Peptide bond
3
1
2
fMet
Phe
fMet
Phe
Phe
fMet
Movement of ribosome
E
E
U
A
C
A
A
A
U
A
C
A
A
A
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
5?
3?
5?
3?
5?
3?
P
A
P
A
P
A
E
4
Thr
fMet
Phe
E
U
A
C
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A
5
Peptide bond
3
1
2
fMet
Phe
fMet
Phe
Phe
fMet
Movement of ribosome
E
E
U
A
C
A
A
A
U
A
C
A
A
A
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
5?
3?
5?
3?
5?
3?
P
A
P
A
P
A
E
4
Thr
fMet
Phe
E
U
A
C
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A
5
fMet
Phe
Thr
Movement of ribosome
E
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A
6
Peptide bond
3
1
2
fMet
Phe
fMet
Phe
Phe
fMet
Movement of ribosome
E
E
U
A
C
A
A
A
U
A
C
A
A
A
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
5?
3?
5?
3?
5?
3?
P
A
P
A
P
A
E
4
Thr
fMet
Phe
E
U
A
C
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A
5
fMet
Phe
Thr
Movement of ribosome
E
U
A
C
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A
Two more cycles
Growing polypeptide
fMet
Phe
Thr
Ala
Ala
Gin
E
C
G
A
C
G
U
A
A
A
U
G
U
A
C
A
U
U
G
G
C
A
C
A
A
G
C
U
G
G
G
3?
5?
P
A